Previous: References
Papers I-VI (Abstracts from
Medline)
Paper I:J Intern Med
1996 Feb;239(2):119-30
Hereditary angio-oedema: new clinical
observations and autoimmune screening, complement and kallikrein-kinin analyses.
Nielsen EW, Gran JT, Straume B, Mellbye
OJ,
Johansen HT, Mollnes TE
Department of Anaesthesiology, Nordland
Central Hospital, Bodo, Norway.
OBJECTIVES. To study clinical and laboratory
manifestations of hereditary angio-oedema (HAE). SUBJECTS. Thirty-three affected members
of a kindred of 63. RESULTS. Oedematous attacks in the skin, mucous membranes and
gastrointestinal tract with fluid displacement were elicited by mental and physical
stress, minor traumas, dental and surgical procedures, eruption of teeth,
tonsillitis, pregnancies, and use of oestrogen-containing pills including menopausal
substitution.
Every adult woman with symptomatic HAE (n = 11) showed symptoms of urinary tract
infections in conjunction with the attacks (P = 0.010), and also experienced more
spontaneous abortions or premature labours (P = 0.037) than healthy relatives. Patients
with HAE of both sexes more frequently reported heartburn or peptic ulcers (P = 0.002).
Rheumatic complaints were reported by 53% of HAE patients and 12% of their unaffected
relatives (P = 0.013), but biochemical screening for 18 autoantibodies and quantitation of
immunoglobulins did not reveal statistically significant differences between the two
groups. C3, prekallikrein, total kininogen, high molecular weight kininogen (HK),
alpha-2-macroglobulin and factor XII were not significantly different in HAE
patients. In contrast, levels of C1-INH and C4 were depressed and cleaved HK increased in patients
compared to unaffected relatives. CONCLUSIONS. HAE manifests in a variety of
ways, and may
influence risk of spontaneous abortions and premature labour.

Paper II: J Immunol
Methods 1994 Aug 1;173(2):245-51
Effect of time, temperature and additives on
a functional assay of C1 inhibitor.
Nielsen EW, Johansen HT, Straume B, Mollnes
TE
Department of Anaesthesiology, Nordland
Central Hospital, Bodo, Norway.
There are different recommendations for the
handling of blood samples for analyses of the kallikrein-kinin or complement system,
respectively. C1 inhibitor (C1-INH) takes a crucial part in both systems. In order to
establish recommendations for blood specimen collection and transport for making the
diagnosis of hereditary angioedema (HAE), the effect of time, temperature and different
additives on C1-INH function and antigen was determined. We used blood samples from
normals and patients suffering from HAE type I. Plasma containing EDTA,
heparin, sodium
citrate or polybrene-EDTA, and serum were assayed after incubations at 4 degrees C or 37
degrees C for 6 or 24 h. In addition, pooled serum was incubated for up to 5 days at room
temperature. A modest decrease in C1-INH function was observed as an effect of
storage-time in samples from normals (p = 0.039) and a substantial decrease was seen for
the HAE patients (p = 0.0002). No significant effect of temperature (4 degrees C or 37
degrees C) was found. Clotting did not reduce C1-INH activity. Plasma containing heparin
or polybrene interfered with the functional assay, yielding falsely high and low
values, respectively. C1-INH functional assay performed within 24 h in serum, EDTA-treated or
citrated plasma discriminated well between HAE patients and normals. This was also the
case for serum kept at room temperature for up to 5 days, although a modest fall in C1-INH
function was seen in the incubation period. For practical purposes we recommend serum as
the sample of choice, preferably received within 48 h.

Paper III: Pediatr Res
1994 Feb;35(2):184-7
C1 inhibitor and diagnosis of hereditary
angioedema in newborns.
Nielsen EW, Johansen HT, Holt J, Mollnes TE
Department of Anaesthesiology, Nordland
Central Hospital, Bodo, Norway.
Symptoms of hereditary angioedema may present
during the child's first years. Attacks may be a particular threat to the narrower airway
of the child. An early diagnosis is most valuable because effective C1 inhibitor (C1
INH)
concentrate is available. We present a reference area for the antigenic and functional
determination of C1 INH by using uncontaminated umbilical cord blood from 80 normal
newborns collected by puncturing vessels in the newly delivered placenta. We examined two
full-term babies (1 and 2) from mothers with hereditary angioedema type I the same
way.
The concentration of C1 INH antigen was determined by radial immunodiffusion. The C1 INH
functional assay was based on the addition of a known quantity of C1s, which enzymatically
splits a chromogenic substrate. The test was performed in the presence of methylamine and
heparin in a kinetic microtiter plate assay. Citrated plasma was used in both
assays. The
data obtained in the 80 cord blood samples (2.5-97.5 percentile) were 0.11-0.22 g/L for C1
INH antigen (adults, 0.15-0.33 g/L) and 47.2-85.9% for C1 INH function (percentage of
adults). In cord blood, baby 1 had an antigenic value of 0.12 g/L (7.5
percentile) and C1
INH function of 61.8% (42 percentile). The corresponding values for baby 2 in cord blood
were less than 0.05 g/L (0.106 g/L < 2.5 percentile) and 34.3% (12.9% < 2.5
percentile). Baby 2 had markedly lower C4 values yet much higher C4 activation products
than baby 1. At 4 mo, baby 1 had an antigenic C1 INH value of 0.24 g/L.

Paper IV: Scand J
Immunol 1995 Dec;42(6):679-85
C3 is activated in hereditary
angioedema,
and C1/C1-inhibitor complexes rise during physical stress in untreated
patients.
Nielsen EW, Johansen HT, Gaudesen O, Osterud
B, Olsen JO, Hogasen K, Hack CE, Mollnes TE
Department of Anaesthesiology, Nordland
Central Hospital, Bodo, Norway.
Seven patients with hereditary angioedema
(HAE) were studied to understand further how physical exercise may induce
attacks. The
most pronounced differences between patients and controls, however, were independent of
the controlled bicycle run (mean values in patients/ controls); C4(g/L): 0.12/0.28 (P =
0.0122); C4bc (AU/ml): 137.0/18.0 (P = 0.0002); C4d (mg/mL): 5.03/2.35 (P = 0.0004); C3bc
(AU/ml): 8.4/6.3 (P = 0.0049); C3a (AU/ml): 11.1/5.6 (P = 0.0102). The ratio C4bc to C4
was 1141 versus 64. Consequently, a substantial part of the low amount of C4 left in HAE
patients consists of activation products, and the authors show for the first time that a
mild but significant activation of C3 occurs in HAE. The two HAE patients treated with
danazol had values of C1-INH function and antigen, C4, and C2 in-between those of normal
and untreated patients, and lower levels of split products from C4 and high molecular
weight kininogen than untreated patients. As a result of the exercise, fibrinolysis
increased significantly in both patients and controls, while C1/C1-INH complexes rose
significantly only in the five HAE patients without treatment when compared to the seven
controls (P = 0.0089). This study thus suggests that complement activation is enhanced in
untreated HAE patients following physical stress.

Paper V: Thromb Haemost
1995 Oct;74(4):1103-6
Factor VIIa in patients with C1-inhibitor
deficiency.
Nielsen EW, Morrissey J, Olsen JO, Osterud B
Department of Anaesthesiology, Nordland
Central Hospital, Bodo, Norway.
In hereditary angioedema (HAE), normal
C1-inhibitor (C1-INH) is low and the contact system activated. Recently, the findings of a
tissue factor mutant selectively deficient in promoting the conversion of FVII to
FVIIa,
but with retained cofactor for FVIIa, made it possible to examine reliably the
pre-existing content of FVIIa in HAE patients. This was of interest as FXIIa
(mainly
inhibited by C1-INH) is able to activate FVII directly. FVIIa in 21 remission HAE patients
were within normal limits but nearly doubled as compared to their 23 normal siblings (p =
0.0017). Cold promoted activation of FVII (CPA) (common clot assay) was displayed in
plasma of all 5 untreated patients (C1-INH function < 35%), but not in plasma of 2
patients treated prophylactically with danazol (C1-INH function about 40%). These results
suggest that there is a minute, yet significant activation of FVII in patients with C1-INH
deficiency.

Paper VI: Scand J
Immunol 1996 Aug;44(2):185-92
Activation of the complement, coagulation,
fibrinolytic and kallikrein-kinin systems during attacks of hereditary angioedema.
Nielsen EW, Johansen HT, Hogasen K,
Wuillemin W, Hack CE, Mollnes TE
Department of Anaesthesiology, Nordland
Central Hospital, Bodo, Norway.
Five patients with hereditary angioedema
(HAE) were studied during attacks and remission as were healthy controls. The high levels
of C1/C1-INH complexes, low C4 and high ratio C4 activation products (C4bc)/C4 also
differed significantly during remission compared to controls. During attacks C4bc/C4
increased (922-2007; P = 0.022, remission versus attacks, median values throughout), C2
and CH50 dropped (111-31%; P = 0.043 and 110-36%; P = 0.016, respectively), TCC (C5b-9)
increased (0.88-1.23 AU/ml; P = 0.028). Cleavage of HK increased to be almost complete
during attacks (20-90%; P = 0.009). While factor XIa/serpin-complexes did not increase, a
more than twofold rise in thrombin/antithrombin-complexes (0.20-0.50 microgram/l; P =
0.009) and in plasmin/alpha-2-antiplasmin-complexes (7.3-17 nmol/l; P = 0.028) was
observed. For the first time cascade activation in HAE was studied simultaneously, and
corroborates that attacks lead to activation of the kallikrein-kinin system, fibrinolysis
and early part of the classical complement pathway. In addition, the authors present novel
data of terminal complement and coagulation activation, the latter apparently not via
FXIa.